oa Annals of the Natal Museum - A revision of Afrotropical Setomima, elucidation of their genealogical relationships and descriptions of other Afrotropical Psychodinae (Diptera: Psychodidae)

Abstract

Using revised morphological terminology, descriptionptions are given of 4 new species of Neoarisemus Botosaneanu & Vaillant, 1 of Tonnoiriella Vaillant, 3 of Setomima Enderlein, and 1 each of Elsahowia Duckhouse, Clogmia Enderlein and Clytocerus Eaton. The following new genus groups are defined: Setomima subgenus Ophryosetomima subg. n. and Mormia Enderlein subgenus Afromormia subg. n. These taxa are placed in tribes, as named by Enderlein, that differ in composition from those recognised by other authors. Setomima is redefined, using a series of new characters. The African species are revised, a key is provided, and the genealogical relationships of the genus are analysed. It is shown that this is a relatively primitive (cladistically basal) genus of Maruinini, outstanding for its structural diversity, and by virtue of its primitive characteristics, linked morphologically with Alepia Enderlein, Neoarisemus and Paratel­matoscopus Satchell, but not necessarily close to them genealogically. Gerobrunettia Quate & Quate is annectant between Setomima and Brunettia Annandale, and Gerobrunettia, Brunettia and Mormia (the Tribe Mormiini) form the apomorph sister group of Setomima and its immediate relatives in the Tribe Maruinini. It appears that the furca of Paramormiini evolved from a 'ball-and-socket' articulation between gonocoxal apodemes and aedeagal apodeme, as seen in Maruinini. The following taxonomic changes are included: Satchelliella Vaillant is a new subjective synonym of Pneumia Enderlein, Krekiella Vaillant and Seoda Enderlein are new Subjective synonyms of Telmatoscopus subg. Xenapates Eaton (as used by Enderlein 1935 1937); Telmatoscopini Vaillant is a new synonym of Paramormiini Enderlein; Brunettiini Vaillant is a new synonym of Mormiini Enderlein; Panimerus subg. Parapanimerus Wagner is a subgenus of Paramormia Enderlein (new combination).